15.1cm25.0cm
Branching Brownian motion with self repulsion
Abstract.
We consider a model of branching Brownian motion with self repulsion. Self-repulsion is introduced via change of measure that penalises particles spending time in an -neighbourhood of each other. We derive a simplified version of the model where only branching events are penalised. This model is almost exactly solvable and we derive a precise description of the particle numbers and branching times. In the limit of weak penalty, an interesting universal time-inhomogeneous branching process emerges. The position of the maximum is governed by a F-KPP type reaction-diffusion equation with a time dependent reaction term.
Key words and phrases:
branching Brownian motion, excluded volume, extreme values, F-KPP equation2000 Mathematics Subject Classification:
60J80, 60G70, 82B441. Introduction
Branching Brownian motion (BBM) [23, 3] can be seen as an elementary model for the evolution of a population of individuals that are subject to birth, death, and motion in space. One of the primary interests in this model was the analysis of the speed of spread of such a population in space, as well as finer properties of the front. Indeed, BBM was investigated form the point of view of extreme value theory over the last 40 year, see, e.g., [9, 19, 12, 13, 5, 6, 7, 4, 14, 8].
As a model for population dynamics, BBM is somewhat unrealistic as it leads to uncontrolled exponential growth of the population size. In fact, in the standard normalisation, the population size grows like , while the population spreads over a volume of order , leading to an unsustainable density of the population. Several variants of the model that resolve this problem have been proposed where, according to some selection rule, offspring is selected to survive in such a way that the total population size stays controlled [11, 21, 15, 20]. Versions where competitive interactions between particles are present were considered, e.g. in [16, 17, 2, 1].
In this paper, we propose a model where the population size is controlled by penalising the fact that particles stay close to each other. Before defining the model precisely, recall that BBM is constructed as follows: start with a single particle which performs a standard Brownian motion in with and continues for a standard exponentially distributed holding time , independent of . At time , the particle splits independently of and into offspring with probability , where , and . In the present paper, we choose the simples option, , all others zero, except in Section 8, where we allow for . These particles continue along independent Brownian paths starting from and are subject to the same splitting rule. And so on. We let denote the number of particles at time and label the particles at time arbitrarily by , and denote by the positions of these particles at that time. For , we let be the position of the ancestor of particle at time . We denote by the law of BBM.
Alternatively, BBM can be constructed as a Gaussian process indexed by a continuous time Galton-Watson tree with mean zero and covariances, conditioned on the Galton-Watson tree, given by
(1.1) |
where is the time of the most recent common ancestor of the particles labeled and in the Galton-Watson tree.
For and for some , we define the penalty function
(1.2) |
(The notation here is not quite consistent, as the labelling of the particles at time is changing with . This can be remedied by using the Ulam-Kesten-Harris labelling of the tree, but maybe this is not necessary here.) We are interested in the law of under the tilted measure given by
(1.3) |
for any Borel set . The function measures the total time when any two particles stay within a distance up to time . This seems to a be reasonable measure for competitive pressure. In a typical realisation of BBM, the density of particles at time will be of order , and hence the -neighbourhood of any particle contains other particles. Thus, for a typical configuration of BBM, . This penalty is most easily avoided by reducing the particle number by not branching. For a particle to not branch up to time has probability , which is far less costly. Reducing the particle density by making the particles move much farther apart would be far more costly.
A simplified model.
Analysing the measure directly seems rather difficult. We suggest an approximation that should share the qualitative features of the full measure. For this we consider a lower bound on . Note that, whenever branching occurs, the offspring start at the same point and thus are all closer than Let us for simplicity take a branching law such that , i.e. only binary branching occurs. Then we can bound
(1.4) |
where is the first time the two Brownian motions that start at the -th branching event are a distance apart. For small , the probability that one of the two branches branches again before the time is of order , so that it will be a good approximation to treat the as independent and having the same distribution as
(1.5) |
Then,
(1.6) |
But (as follows from Theorem 5.35 and Proposition 7.48 in [22]),
(1.7) |
which for small behaves like . Note that we also have, by Jensen’s inequality, that
(1.8) |
One might think that the approximate model is a poor substitute for the full model, since it ignores the repulsion of particles after the time that they first separate. However, as we will see shortly, already suppresses branching so much that the total number of particles will stay finite for any time. Hence we can expect that these finitely many particles can remain separate rather easily and that the remaining effect of will be relatively mild.
Outline.
The remainder of this paper is organised as follows. In Section 2 we derive exact formulas for the partition function, the particle number, and the first branching time in the simplified model. In Section 3 we introduce the notion of quasi-Markovian Galton-Watson trees. In Section 4 we show that the branching times in the simplified model are given by such a tree. In Section 5, we consider the limit when and derive a universal asymptotic model, which is a specific quasi-Markovian Galton-Watson tree. In Section 6, we consider the position of the maximal particle and show that its distribution is governed by a F-KPP equation with time dependent reaction term and analyse the behaviour of its solutions. We discuss the relation of the approximate model to the full model in Section 7. In Section 8, we briefly look at the case when . In this case, the process dies out and we derive the rate at which the number of particles tends to zero.
2. Partition function, particle numbers, and first branching time
2.1. The partition function
The first object we consider is the normalising factor or partition function
(2.1) |
Lemma 2.1.
Let be the solution of the ordinary differential equation
(2.2) |
with initial condition . Then .
Proof.
A first inspection of Eq. (2.2) shows why the cases and are vastly different.
Equation (2.2) has the two fix points and . Here is stable and is unstable. Hence all solutions with initial condition will converge to , while solutions with will tend to infinity. Only the special initial condition will lead to the constant solution. Since we start with the initial condition , if and hence , we get this special constant solution, while for , the solution will tend to zero. In fact, we can solve (2.2) exactly. To do this it is convenient to define
(2.5) |
Then solves
(2.6) |
also with initial condition . Dividing both sides by , this can be written as
(2.7) |
which can be integrated to give
(2.8) |
or
(2.9) |
and
(2.10) |
Using the initial condition , we get
(2.11) |
Thus, provided ,
(2.12) |
2.2. Particle numbers
From the formula for the partition function we can readily infer the mean number of particles at time , nramely,
(2.13) |
and for this converges to . For small , this in turn behaves like .
In fact, we can even compute the distribution of the number of particles at times . To do so, we want to compute the Laplace (Fourier) transforms
(2.14) |
The denominator has already been calculated. For the numerator we write
(2.15) | |||||
where are the number of particles at time that have particle as common ancerstor at time . Using the independence properties, this equals
(2.16) |
Dividing by , we arrive at
(2.17) |
From this exact formula we can derive various special cases.
Theorem 2.2.
-
(i)
Under the measure , the number of particles at time is geometrically distributed with parameter . In particular, the number of particles converges, as , to a geometric random variable with parameter .
-
(ii)
As , the number of particles at time , for all , converges in distribution to a geometric random variable with parameter
.
Proof.
Inserting into (2.17) we get that
(2.18) |
which is the Laplace transform of the geometric distribution with parameter . This implies (i). Similarly, with , and
(2.19) |
If we now take , we get
(2.20) |
which is the Laplace transform of the geometric distribution with parameter .
∎
Remark.
Note that, for fixed , taking the limit , we get unsurprisingly , indicating that there is just one particle.
We see that the mean number of particles ranges from (as ), (for ), to (for ). Note that if , is geometric with parameter , which corresponds to BBM with binary branching.
2.3. Distribution of the first branching time
We have seen so far that the repulsion strongly suppresses the number of branchings. The first branching time is then
(2.21) |
Theorem 2.3.
The distribution of the first branching time under is given by
(2.22) |
Proof.
Note that after the first branching, there will be two independent BBMs that run for the remaining time and that are subject to the same penalty as before. In particular, given , the total particle number is equal to the sum of the number of particles in these two branches,
(2.23) |
where are the particles in the two branches that split at time . Denote by the unnormalised mass of paths that branch before time , i.e. set
(2.24) |
We get
(2.25) | |||||
Since is known, this is an explicit formula, namely
(2.26) | |||||
Since , (2.22) follows. ∎
Remark.
Note that, for fixed, converges, as , to
(2.27) |
Note further that and therefore
(2.28) |
3. Quasi-Markovian time-inhomogeneous Galton-Watson trees
In this section we introduce a class of models that are continuous-time version of Galton-Watson processes that are time-inhomogeneous and that in general are not Markov, but have an underlying discrete-time Markov property. These processes emerge in the models introduced above.
We start with discrete time trees and we introduce the usual Ulam-Harris labelling.
Let us define the set of (infinite) multi-indices
(3.1) |
and let denote the subset of multi-indices that contain only finitely many entries that are different from zero. Ignoring leading zeros, we see that
(3.2) |
where is either the empty multi-index or the multi-index containing only zeros. A discrete-time tree is then identified by a consistent sequence of sets of multi-indices, at time as follows.
-
•
.
-
•
If then if , where
(3.3)
We can relate the assignment of labels in a backwards consistent fashion as follows. For , we define the function , through
(3.4) |
Clearly, if and , then . This allows to define the boundary of the tree at infinity as follows:
(3.5) |
We also want to be able to consider a branch of a tree as an entire new tree. For this we use the notation if .
Given a discrete-time tree, we can turn it into a continuous-time tree by assigning waiting times to each vertex, resp. to each multi-index in the tree. E.g., in the case of the standard continuous-time Galton-Watson tree, we simply assign standard, iid, exponential random variables, , to each vertex, resp. multi-index. Note that we choose the notation in such a way that we think of as an element of the boundary of the tree, and is the waiting time attached to the vertex labelled (in the -th generation). This time represents the waiting time from the birth of this branch to its next branching. This assigns a total time, for the branching of a multi-index at discrete time , as
(3.6) |
where is an initial time associated to the root of the tree and is the time of the first branching of the root of the tree. We denote by the -algebra generated by the branching times of the first generations of the tree, i.e.
(3.7) |
We need to define further -algebras that correspond to events that take place in sub-trees. For a given multi-index , define the set of multi-indices that coincide with in the first entries,
(3.8) |
Naturally, this is the subtree that branches off the branch in the -th generation. Next, we define the -algebra generated by the times in these subtrees,
(3.9) |
Definition 3.1.
A set is called normal, if it is of the form
(3.10) |
or if
(3.11) |
where the two events are normal. We say that a normal event has finite horizon, if there exists a such that .
Definition 3.2.
We say that the assignment of branching times is quasi-Markov (with time horizon ), if there is a family of probability measures, on and a family of probability measures on that have the following property. For any event which is of the form
(3.12) |
where , for all ,
(3.13) |
Lemma 3.3.
The measures on the -algebra generated by the normal events with finite horizon in are uniquely determined by the family of measures . is the law of conditioned on .
Proof.
From (3.13) it follows by simple iteration that the measure of any normal event of finite horizon is expressed uniquely in term of . Noting further that the set of finite horizon events is intersection stable, the assertion follows from Dynkin’s theorem. ∎
The total tree at continuous time is then described as follows:
-
(i)
The branches of the tree alive are
(3.14) -
(ii)
The entire tree up to time is the set
(3.15)
Note that both sets are empty if . It is a bit cumbersome to write, but the distribution of the set together with the lengths of all branches can be written down explicitly in terms of the laws and the branching laws of the underlying discrete-time tree.
4. The simplified model as quasi-Markov Galton-Watson tree
We return to the approximate model defined in Section 1. For simplicity, we keep the assumption that the underlying tree is binary. We first show that the branching times under the law define a quasi-Markov Galton-Watson tree.
Lemma 4.1.
The branching times of the simplified model under the law are quasi-Markov, where is the marginal distribution of with that is absolutely continuous w.r.t. Lebesgue mesure with density
(4.1) |
namely,
(4.2) |
Proof.
Let . We now have
(4.3) |
where the are the particle numbers in the four respective branches of the tree. In analogy to (2.25), we obtain
(4.4) |
where we used the independence the events in the two branches under the original BBM measure and the definition of . This concludes the simple proof. ∎
5. The limit
We have seen that a penalty with fixed and enforces that only a finite number of branchings take place, even if we let tend to infinity. To get more interesting results, we consider now the case when depends on such that as . In fact, we will see that a rather interesting limiting model arises in this setting. Clearly, in this case is a good approximation.
We first look at the partition function.
Lemma 5.1.
Assume that . Then
(5.1) |
Proof.
We just use the explicit form of given in (2.11). This gives
(5.2) | |||||
which implies the statement of the lemma. ∎
From Theorem 2.2 we derive the asymptotics of the particle number.
Theorem 5.2.
Assume that , but , as . Then:
-
(i)
The number of particles at time times , , converges in distribution to an exponential random variable with parameter .
-
(ii)
For any , the number of particles at time converges in distribution to a geometric distribution with parameter .
-
(iii)
If but , the number of particles at time divided by converges in distribution to an exponential random variable with parameter .
Proof.
The next theorem gives the asymtotics of the first branching time.
Theorem 5.3.
Let be as in Theorem 5.2. Then, for any ,
(5.3) |
Proof.
5.1. The limiting Quasi-Markov Galton-Watson tree
Theorem 5.3 also suggests to define
(5.5) |
should be thought of as the position of the first branching seen from the standard position .
To derive the asymptotics of the consecutive branching times, we just have to look at
(5.6) |
For this we have from the previous computation
(5.7) |
Recall that, e.g. , where is a finite random variable, so that we will have that, in general, , so that the right-hand side of (5.7) will converge to some non-trivial distribution.
The asymptotic results above suggest to consider the branching times of the process in the limit , , around the time . We have seen that the time of the first branching shifted by this value converges in distribution to a random variable with distribution function (which is supported on ).
This suggests to define a limiting model as a quasi-Markov Galton-Watson tree with the measures
(5.8) |
This gives, in particular, for the first branching time,
(5.9) |
We have to choose to match this with the known asymptotics of the first branching time, see (5.3). It turns out that
(5.10) |
for all . So the picture is that we start the process at and the first branching time is infinitely far in the future and occurs at a finite random time distributed according to (5.10). The density of this distribution is . In particular, it has mean zero and variance .
We have the following result.
Theorem 5.4.
Assume that and , as . Then, for any and events ,
(5.11) |
where is the law of the limiting model.
Proof.
All we need to show is that the measures converge. But this follows form the computations indicated above. ∎

Note further, as long as is negative, the distribution of is concentrated around , while as become positive and large, the distribution tends to a standard exponential distribution. In fact,
(5.12) |
Clearly this converges to , as and behaves like , as tends to .
6. The distribution of the front
An obvious first question is the distribution of the maximum of BBM under the law . We define, for any ,
(6.1) |
Then
(6.2) |
Note that we use the choice to be closer to the usual formulation of the F-KPP equation.
Interestingly, solves a time-dependent version of the F-KPP equation.
Lemma 6.1.
Proof.
In complete analogy to the derivation of the F-KPP equation (see, e.g. [8]), satisfies the recursive equation
(6.4) |
where is the probability that a single Brownian motion at time is smaller than . Letting , we can write this as
(6.5) |
Note that is the Green function for the differential operator . and so is the mild formulation of the partial differential equation
(6.6) |
with initial condition . This equation is the F-KPP equation if , i.e. if , and looks similar to it in general. Hence,
(6.7) | |||||
where we used the explicit form of from (2.11). ∎
Note that (6.3) is a time-dependent version of the F-KPP equation, where the non-linear term is modulated down over time. Time dependent F-KPP equations have been studied in the past, see, e.g. [25, 18, 24], but we did not find this specific example in the literature. For small , (6.3) becomes
(6.8) |
For future use, note that (6.8) is a special case of a class of F-KPP equations of the form
(6.9) |
where . We will be interested in the case when is bounded, monotone decreasing, and integrable.
The key tool for analysing solutions of (6.3) is the Feynman-Kac representation for , see Bramson [10].
Lemma 6.2.
If is a solution of the equation (6.9) with initial condition , then satisfies
(6.10) |
where is Brownian motion starting in .
The strategy to exploit this representation used by Bramson is to use a priori bounds on in the right-hand side of the equation in order to get sharp upper and lower bounds. Here we want to do the same, but we need to take into account the specifics of the function . Going back to the specific case (6.8), remains close to for a fairly long time (), and then decays exponentially with rate to zero. Therefore, we expect that initially, the solution will behave like that of the F-KPP equation and approach a travelling wave solution. As time goes on, the wave slows down and comes essentially to a halt. Finally, as time goes on we see a pure diffusion. We will deal differently with these three regimes. We begin with the initial phase when .
Lemma 6.3.
Assume that is non-increasing and bounded by one and zero from above and below. Define . Then
(6.11) |
Proof.
can be computed explicitly for , namely
(6.13) |
Notice that
(6.14) |
Define, for ,
(6.15) |
Obviously,
(6.16) |
so
(6.17) |
Finally, .
In the limit , we get
(6.18) |
(6.19) |
and
(6.20) |
We see that, as , . This allows us to deduce the precise behaviour of the solution at this time via Bramson’s results.
Lemma 6.4.
If satisfies (6.3) with Heaviside initial condition Then, as ,
(6.21) |
where is a travelling wave of the F-KPP equation with speed
(6.22) |
and .
Next we look at the behaviour of the solution for times when .
Lemma 6.5.
Let solve (6.9) and be integrable. Define, for , by
(6.23) |
Then, for ,
(6.24) |
where is Brownian motion started in .
Proof.
Note that in our case, is determined by
(6.27) |
But
Now set . Then
(6.29) |
Hence
(6.30) |
for small . In particular, we have that
(6.31) |
for small and .
What is left to do is to control the evolution of the solution between time and . The Feynman-Kac representation, for ,
(6.32) |
To start, the following bounds are straightforward from the Feynman-Kac representation and the fact that .
Lemma 6.6.
With the notation above, for ,
(6.33) |
where
(6.34) |
and .
Remark.
We expect the upper bound to be closer to the correct answer.
We now combine all estimates. This gives, for ,
(6.35) |
If we choose, e.g. , we see that the upper and lower bounds only differ by a factor .
The expectation over can be bounded using the known tail estimates (see [10] and [14]),
(6.36) | |||||
(6.37) |
However, the resulting expressions are not very nice and not very precise, so we leave their computation to the interested reader.
We conclude this chapter by summarising the behaviour of the solution as a function of when .
Theorem 6.7.
This picture corresponds to the geometric picture we have established in the preceding sections, in a sort of time reversed way: the diffusive behaviour at large times corresponds to the Brownian morion up to the time of the first branching , the travelling wave behaviour at times up to corresponds to the almost freely branching at the late times after , and the finite time interval between and , when the travelling waves comes to a halt corresponds to the first branching steps that are asymptotically described by the limiting quasi-Markov Galton-Watson tree described in Section 5.
7. Comparison to the full model
We will show that in the original model, with the interaction given by (see Eq. (1.2)), behaves similarly to the simplified model. In particular, the first branching happens at least as late as in that model.
Lemma 7.1.
Let be the first branching time. Then
(7.1) |
For and , this behaves as
(7.2) |
and so
(7.3) |
Proof.
Set
(7.4) |
Then
(7.5) |
where we simply bounded the denominator by the probability that there is no branching up to time . Inserting the explicit form of gives (7.1). The asymptotic formulae for small are straightforward. ∎
One can improve the bound above as follows. Instead of bounding the denominator just by the probability that there is no branching up to time , we can bound it by no branching up to time and then bound the interaction of the remaining piece uniformly by
(7.6) |
Hence the denominator becomes
(7.7) |
Now
(7.8) | |||||
Since is a positive martingale, by Doob’s maximum inequality we have that, for ,
(7.9) |
Moreover, , and hence
(7.10) |
Finally, we make the close to optimal choice , which yields
(7.11) |
Thus, choosing , (7.3) improves to
(7.12) |
Hence,
(7.13) |
This is still not perfect for small , but it seems very hard to improve the bound on the denominator much more. Improvement would need to come from a matching bound in the numerator.
8. The case
The behaviour of the model is very different if particles are allowed to die. In that case, the process will die out almost surely. However, it is still interesting to see how exactly this happens. To simplify things, we assume in the sequel and . Note first that the approximate penalty function changes slightly, since now the number of branching events is no longer related to the number of particles. Let us introduce the two numbers and as the number of (binary) branchings and deaths, resp. that occurred up to time . Clearly, . We then have
(8.1) |
We consider first the partition function function . The analog of Lemma 2.1 is as follows.
Lemma 8.1.
Let be the solution of the ordinary differential equation
(8.2) |
with initial condition . Then .
Proof.
We proceed as in the proof of Lemma 2.1. Since now the first event could be either a branching (with probability ) or a death (with probability ), we get the recursion
(8.3) |
Differentiating yields the asserted claim. ∎
The presence of the term eliminates the fixpoint in equation (8.2). In fact, (8.2) has the two fixpoints
(8.4) |
Note that for , this simplifies to
(8.5) |
which is and . In that case we clearly have in all cases.
If , but (i.e. , but small), we can expand
(8.6) |
In particular, the smaller fixpoint is
(8.7) |
Now set . Then satisfies the differential equation
(8.8) |
with initial condition . We can solve this equation as in the case . Define
(8.9) |
Then
(8.10) |
which has the solution
(8.11) |
Hence
(8.12) |
Note that in the case , this is just one, while otherwise, it decays exponentially to zero, so that , indicating that the number of branchings in the process remains finite.
Next let us consider the generating function of the particle number,
(8.13) |
We readily see that this function satisfies the equation
(8.14) |
This implies the differential equation
(8.15) |
with initial condition . Thus, and differ only in the initial conditions. it is therefore easy to see that
(8.16) |
From this expression we can compute, e.g., the expected number of particles at time under the measure ,
(8.17) |
which reads
(8.18) |
For , this behaves to leading order, provided , like
(8.19) |
This implies that the process dies out exponentially fast unless the death rate is zero. This does not come, of course, as a surprise.
Alternative computation of .
Instead of passing through the generating function for , we can also proceed by deriving a direct recursion for . To do so, define the un-normalised expectation
(8.20) |
Clearly we have
(8.21) |
where we used that in the case when the first event is a death at time , will be zero, while in the case of a birth, , where are independent copies. This implies the differential equation
(8.22) |
The solution of this can be written directly as
(8.23) |
Since is explicit, one can verify that this gives the same answer as (8.18).
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